Pantherophis quadrivittatus

Familia: Colubridae
Subfamilia: Colubrinae
Genus: Pantherophis
Species: Pantherophis quadrivittatus

Pantherophis quadrivittatus (Holbrook, 1836)

• Holotype: ANSP 3773.
• Type locality: “South Carolina”, restricted to “Charleston, South Carolina” by Schmidt (1953: 198).

• Coluber quadrivittatus Holbrook, 1836: 113 [original combination]
• Elaphis quadri-vittatus — Duméril, Bibron & Duméril, 1854: 265 [subsequent combination]
• Elaphe obsoleta quadrivittata — Neill, 1949: 4 [subsequent combination]
• Pantherophis quadrivittatus — Hillis & Wüster, 2021: 52 [subsequent combination]

• Elaphe quadrivittata deckerti Brady, 1932: 5
  • Holotype: USNM 84295, ♂, collected by R.F. Deckert in June 1931.
  • Type locality: “Lower Matecumbe Key, Monroe Co. Florida”.

• Elaphe williamsi Barbour & Carr, 1940: 340
  • Holotype: MCZ 45705.
  • Type locality: “near Lebanon, Levy County, Florida”.

• Elaphe quadrivittata parallela Barbour & Engels, 1942: 103
  • Holotype: MCZ 46468.
  • Type locality: “Sam Windsor's Lump, [Shackleford Banks, Carteret County,] North Carolina”.

• Elaphe obsoleta rossalleni Neill, 1949: 1
  • Holotype: UF 16264 (formerly WTN 18881), adult ♂, collected by W.T. Neill on 11 July 1947.
  • Type locality: “22 miles northwest of Fort Lauderdale, Broward County, Florida (near junction of State Highways 25 and 84)”.

• United States: Virginia, North Carolina, South Carolina, Georgia and Florida.

• Holbrook, J.E. 1836–1840. North American Herpetology; or, A description of the reptiles inhabiting the United States. J. Dobson: Philadelphia. Volume I: 120 pp. + XXIII pls. BHL; Volume II: 125 pp. + XXX pls. BHL; Volume III: 122 pp. + XXX pls. BHL; Volume IV: 126 pp. + XXVIII pls. BHL Reference page. 
• Brady, M.K. 1932. A new snake from Florida. Proceedings of the Biological Society of Washington 45: 5–7. BHL Reference page. 
• Barbour, T. & Carr, A.F. 1940. Notes on Elaphe and a new species. Occasional papers of the Boston Society of Natural History 8: 337–342. Reference page. 
• Barbour, T. & Engels, W.L. 1942. Two interesting new snakes. Proceedings of the New England Zoölogical Club 20: 101–104. Reference page. 
• Neill, W.T. 1949. A New Subspecies of Rat Snake (Genus Elaphe), and Notes on Related Forms. Herpetologica 5(Second Supplement): 1–11. JSTOR Reference page. 
• Schmidt, K.P. 1953. A Check List of North American Amphibians and Reptiles. 6th edition. University of Chicago Press: Chicago. viii + 280 pp. Reference page. 
• Hillis, D.M. & Wüster, W. 2021. Taxonomy and Nomenclature of the Pantherophis obsoletus Complex. Herpetological Review 52(1): 51–52. Reference page. 

• Burbrink, F.T., Gehara, M., McKelvy, A.D. & Myers, E.A. 2021. Resolving spatial complexities of hybridization in the context of the gray zone of speciation in North American ratsnakes (Pantherophis obsoletus complex). Evolution 75(2): 260–277. DOI: 10.1111/evo.14141 Reference page. 
• Burbrink, F.T., Pyron, R.A., Gehara, M., McKelvy, A.D. & Myers, E.A. 2021. The corrected taxonomic history of the North American ratsnakes (Pantherophis obsoletus complex). Herpetological Review 52(3): 537–547. Reference page. 

For more multimedia, look at Pantherophis quadrivittatus on Wikimedia Commons.