1 Key Laboratory of Alpine Ecology, Institute of Tibetan Plateau Research, Chinese Academy of Sciences, Beijing 100101, China.
2 State Key Laboratory of Urban and Regional Ecology, Research Center for Eco-environmental Sciences, Chinese Academy of Sciences, Beijing 100085, China; State Key Laboratory of Earth Surface Processes and Resource Ecology, Faculty of Geographical Science, Beijing Normal University, Beijing 100875, China. Electronic address: bfu@rcees.ac.cn.
3 State Key Laboratory of Earth Surface Processes and Resource Ecology, Faculty of Geographical Science, Beijing Normal University, Beijing 100875, China.
4 School of Environment, Beijing Normal University, Beijing 100875, China.
5 State Key Laboratory of Urban and Regional Ecology, Research Center for Eco-environmental Sciences, Chinese Academy of Sciences, Beijing 100085, China.
6 Key Laboratory of Restoration Ecology for Cold Regions in Qinghai, Northwest Institute of Plateau Biology, Chinese Academy of Sciences, Xining 810008, China.
7 College of Grassland Science and Technology, China Agricultural University, Beijing 100193, China.
8 Key Laboratory of Aquatic Botany and Watershed Ecology, Wuhan Botanical Garden, Chinese Academy of Sciences, Wuhan 430074, China.
9 Qilian Mountain Nature Reserve Bureau of Qinghai Province, Xining 810008, China.